Acta Biologica Szegediensis


Acta Biologica Szegediensis (ISSN 1588-385X print form; ISSN 1588-4082 online form), a member of the Acta Universitatis Szegediensis family of scientific journals (ISSN 0563-0592), is published yearly by the University of Szeged. Acta Biologica Szegediensis covers the growth areas of modern biology and publishes original research articles and reviews, involving, but not restricted to, the fields of anatomy, embryology and histology, anthropology, biochemistry, biophysics, biotechnology, botany and plant physiology, all areas of clinical sciences, conservation biology, ecology, genetics, microbiology, molecular biology, neurosciences, paleontology, pharmacology, physiology and pathophysiology, and zoology. Occasionally, Acta Biologica Szegediensis will publish symposium materials. Acta Biologica Szegediensis particularly encourages young investigators and clinicians to submit novel results of interest.

Editor-in-Chief: Károly Gulya

Senior Editors: Dénes Budai (Cell Physiology) Julius Gy. Papp (Pharmacology) István Raskó (Genetics)

Editorial Board: Magdolna Ábrahám-Gulyás (Biochemistry) Mihály Boros (Experimental Surgery) László Erdei (Plant Physiology) Gyula Farkas (Anthropology) László Gallé (Ecology) Zoltán Janka (Psychiatry) Csaba Vágvölgyi (Microbiology) Kornél Kovács (Biotechnology) János Lonovics (Internal Medicine)

Technical Editors: Lidia Bakota, Tamás Mikola

Péter Maróti (Biophysics)

Péter Maróy (Genetics)

Erzsébet Mihalik (Botany)

András Mihály (Anatomy, Embryology, Histology)

Attila Pál (Obstetrics and Gynecology)

Aurél J. Simonka (Traumatology, Surgery)

József Toldi (Comparative Physiology)

László Vécsei (Neurology)

László Vígh (Biochemistry)

Submission of manuscripts

Manuscripts should be prepared in accordance with the Instructions to Authors published in each issue, also available at, and submitted to:

Károly Gulya

Acta Biologica Szegediensis, Editorial Office

Department of Zoology and Cell Biology

University of Szeged

2 Egyetem u., POB 659, H-6722 Szeged, Hungary

Phone: 36 (62) 544-048, fax: 36 (62) 544-049


Correspondence relating to the status of the manuscripts, proofs, publication, reprints and advertising should be sent to:

Lidia Bakota

Acta Biologica Szegediensis, Editorial Office

Department of Zoology and Cell Biology

University of Szeged

2 Egyetem u., POB 659, H-6722 Szeged, Hungary

Phone: 36 (62) 544-125, fax: 36 (62) 544-049



Acta Biologica Szegediensis is published yearly in four issues per volume. All subscriptions relate to the calendar year and must be pre-paid. The annual subscription rate is currently 50 USD and includes air mail delivery and handling.

Acta Biologica Szegediensis is indexed in BIOSIS Database, EMBASE, Excerpta Medica, Elsevier BIOBASE (Current Awareness in Biological Sciences) and Zoological Record.

The Table of Contents for the current issue and those for previous issues can be found at

Instructions to Authors Submission of manuscripts

Submission of a manuscript to Acta Biologica Szegediensis automatically involves the assurance that it has not been published and will not be published elsewhere in the same form. Manuscripts should be written in English. Since poorly-written material will not be considered for publication, authors are encouraged to have their manuscripts corrected for language and usage by a trusted expert. There are no explicit length limitations: a normal research article will occupy 4-6 printed pages; reviews might be considerably longer. Authors should submit three sets of the complete manuscript and illustrations, together with a computer disk containing an electronic version of their manuscript. The elctronic file is considered the final material. Both Macintosh and PC versions will be accepted. The disk should be labeled with the date, the first author's name, the file name of the manuscript and the software, disk format and hardware used. Acta Biologica Szegediensis will not return copies of submitted manuscripts and figures. Requests to return original figures will be honored as a courtesy, but cannot be guaranteed. If instructions are not followed, authors will be asked to retype their manuscripts.

Manuscript format

Only good-quality laser printouts will be accepted. All pages should be printed with full double spacing, 2.5 cm margins, and a nonjustified right margin. A standard 12 point typeface (e.g. Times, Helvetica or Courier) should be used throughout the manuscript, with symbol font for Greek letters. Boldface, italics or underlined text should not be used anywhere in the manuscript. Footnotes are not permitted. Each page should be numbered at the bottom as follows:

Page 1. Title page: Complete title, first name, middle initial, last name of each author; where the work was done (authors' initials in parentheses if necessary); mailing address, phone, fax, and e-mail of the corresponding author; a running title of no more than 48 characters and spaces.

Page 2. Abstract: no more than 200 words, followed by 4-6 key words.

Beginning on page 3: Introduction, Materials and Methods, Results, Discussion, Acknowledgments, References, Figure Legends, Tables. Each section should be begun on a new page.

The manufacturer's name and location should be given in parentheses for reagents and instruments. Sources for all antibodies and nucleotide sequences should be indicated. Customary abbreviations in common use need not be defined in the text (e.g. DNA or ATP). Other abbreviations should be defined the first time that they are used. Quantitative results must be presented as graphs or tables and supported by appropriate experimental design and statistical tests. Only SI units may be used. For studies that involve animals or human subjects, the institutional, national or international guidelines that were followed should be indicated.


Only work that has been published or is in the press may be referred to. Personal communications should be acknowledged in the text and accompanied by written permission. In the text, references should be cited by name and year, e.g. Bloom (1983) or (Schwarz-Sommer et al. 1990) or (Maxam and Gilbert 1977). In the References, references should be listed alphabetically by first authors (including all coauthors) and chronologically for a given author (beginning with the most recent date of publication). Where the same author has more than one publication in a year, lower case letters should be used (e.g. 1999a, 1999b, etc.). Periods should not be used after authors' initials or abbreviated journal titles (e.g. Acta Biologica Szegediensis should be cited as Acta Biol Szeged). Inclusive page numbers should be used. Examples:

Bloom FE (1983) The endorphins: a growing family of pharmacologically pertinent peptides. Annu Rev Pharmacol Toxicol 23:151-170.

Coons AH (1978) Fluorescent antibody methods. In Danielli JF, ed., General Cytochemical Methods. Academic Press, New York, 399-422.

Maxam AM, Gilbert WA (1977) A new method for sequencing DNA. Proc Natl Acad Sci USA 74:560-564.

Monod J, Changeux J-P, Jacob F (1963) Allosteric proteins and cellular control systems. J Mol Biol 6:306-329.

Schwarz-Sommer Z, Huijser P, Nacken W, Saedler H, Sommer H (1990) Genetic control of flower development by homeotic genes in Antirrhinum majus. Science 250:931-936.


Three complete sets, including a high-quality "original" for publication, must be submitted with the manuscript. The back of each figure or composite plate should be labeled in soft lead pencil, indicating the orientation, the figure number, and the first author's name. The back of the best set should be marked "use for reproduction" or "original". Authors are encouraged to submit digital images of photographs, line drawings or graphs for printing. Most major image editing and drawing/illustrator computer software files (both Macintosh and PC) in TIFF or EPS formats are acceptable. It is particularly important that adequate resolution (at least 300 dpi, preferably 600 dpi) is used in making the original image.

Figure legends

Figures should be numbered consecutively with Arabic numerals. Material in the text should not be duplicated and methods should not be described. The size of scale bars should be indicated when appropriate. The first figure in the text should be referred to as Fig. 1, and so on.


Tables should be numbered consecutively with Arabic numerals. A brief title should be included above the table. Each table should be printed double spaced, without vertical or horizontal lines, and on a separate sheet. Material in text should not be duplicated and methods should not be described. The first table in the text should be referred to as Table 1, and so on.